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California Oak Galls
Wasps in the family Cynipidae are gall inducers on plants, or inquilines of gall inducers
(living inside the galls made by other cynipids). The majority of species occur on oak
trees (Quercus sp.). Galls range in size from 1 mm to 6 cm or more and come in many shapes
and colors, as depicted on this page. Different cynipid species occur on specific oak species,
and the galls themselves occur on specific locations on the trees (leaves, stems, buds, roots,
etc.) depending on the species and which generation it is. Some species have two generations
per year, a sexual and an asexual (female only) generation, each producing completely different
types of galls on different parts of the host. The adult wasps are generally around 1-4mm in
size. The wasps support a community of other insects that feed on them, their parasites, and
their galls.
Regarding the generic names I'm using on this web site, please read this page.
See also the undescribed or unknown oak gall page.
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White oak group:
Quercus berberidifolia (inland scrub oak),
Q. cornelius-mulleri (Muller's oak; desert scrub oak),
Q. douglasii (blue oak),
Q. dumosa (Nuttall's scrub oak)
Q. durata (leather oak),
Q. engelmannii (Engelmann oak),
Q. garryana (Oregon oak),
Q. john-tuckeri (Tucker's oak),
Q. lobata (valley oak):
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Andricus albobalani
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Andricus brunneus
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Andricus confertus
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Andricus coortus
(syn: Callirhytis coortus)
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Andricus chrysolepidicola
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Andricus opertus
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Andricus quercuscalifornicus
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Andricus wiltzae
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Antron douglasii
Asexual, fall generation
(syn: Cynips douglasii)
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Antron douglasii
Bisexual, spring generation
(syn: Cynips douglasii)
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Antron quercusechinus
(syn: Cynips quercusechinus)
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Besbicus conspicuus
(syn: Cynips conspicuus)
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Besbicus heldae
(syn: Cynips heldae)
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Besbicus maculosus
(syn: Cynips maculosus)
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Besbicus mirabilis
(syn: Cynips mirabilis)
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Besbicus multipunctatus
(syn: Cynips multipunctata)
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Burnettweldia conalis
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Burnettweldia corallina
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Burnettweldia plumbella
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Burnettweldia washingtonensis
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Disholcaspis canescens
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Disholcaspis eldoradensis
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Disholcaspis mamillana
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Disholcaspis mellifica
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Disholcaspis prehensa
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Disholcaspis simulata
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Druon fullawayi
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Feron atrimentum
Asexual, fall generation
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Feron atrimentum
Bisexual, spring generation
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Feron bakkeri
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Feron californicum
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Feron crystallinum
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Feron discularis
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Feron gigas
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Feron izabellae
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Feron kingi
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Feron parmula
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Feron pattersonae
Asexual, fall generation
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Feron pattersonae
Bisexual, spring generation
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Feron stellare
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Feron stellulum
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Feron tubifaciens
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Neuroterus fragilis
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Neuroterus saltatorius
Asexual, fall generation
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Neuroterus saltatorius
Bisexual, spring generation
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Neuroterus washingtonensis
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Phylloteras cupella
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Sphaeroteras trimaculosum
(syn: Atrusca trimaculosa)
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Xanthoteras clavuloides
(syn: Atrusca clavuloides)
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Xanthoteras teres
(syn: Trigonapsis teres)
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Red (black) oak group: Quercus agrifolia (coast live oak),
Q. kelloggii (California black oak),
Q. wislizeni (interior live oak):
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Amphibolips quercuspomiformis
Asexual, summer generation
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Amphibolips quercuspomiformis
Bisexual, spring generation
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Callirhytis apicalis
(syn: Andricus apicalis)
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Callirhytis carmelensis
(syn: Andricus carmelensis)
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Callirhytis congregata
(syn: Andricus congregata)
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Callirhytis eldoradensis
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Callirhytis perdens
(syn: Andricus perdens)
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Callirhytis perfoveata
(syn: Andricus perfoveata)
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Callirhytis quercusagrifoliae
Asexual, summer generation
(syn: Andricus quercusagrifoliae)
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Callirhytis quercussuttoni
(syn: Andricus quercussuttoni)
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Callirhytis serricornis
Asexual, summer generation
(syn: Andricus serricornis)
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Callirhytis serricornis
Sexual, spring generation
(syn: Andricus serricornis)
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Dryocosmus dubiosus
Asexual, fall generation
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Dryocosmus dubiosus
Bisexual, spring generation
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Dryocosmus minusculus
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Melikaiella flora
Asexual, summer generation
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Melikaiella flora
Bisexual, spring generation
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Intermediate or golden oak group:
Quercus chrysolepis (canyon live oak),
Q. palmeri (Palmer's oak),
Q. vacciniifolia (huckleberry oak):
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Andricus chrysobalani
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Andricus projectus
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Disholandricus chrysolepidis
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Disholandricus lasius
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Disholandricus reniformis
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Disholandricus truckeensis
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Dryocosmus asymmetricus
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Grahamstoneia humboldti
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Heteroecus dasydactyli
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Heteroecus flavens
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Heteroecus lyoni
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Heteroecus melanoderma
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Heteroecus malus
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Heteroecus pacificus
Asexual, fall generation
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Heteroecus pacificus
Bisexual, spring generation
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Heteroecus sanctaeclarae
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Heteroecus sp.
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Holocynips hartmani
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Paracraspis guadaloupensis
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Paracraspis insolens
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Paracraspis patelloides
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Protobalandricus spectabilis
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Trichoteras burnetti
(syn: Andricus burnetti)
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Trichoteras coquilletti
(syn: Andricus coquilletti)
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Trichoteras frondeum
(syn: Andricus frondeum)
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Trichoteras perfulvum
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Trichoteras rotundula
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Trichoteras vacciniifoliae
(syn: Andricus vacciniifoliae)
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Non-native oaks growing in California:
Quercus virginiana (southern live oak)
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Andricus quercuslanigera
This gall has been seen on Q. virginiana in Manteca, Stockton, Brentwood, and Sacramento. It has also been seen in Los Angeles County.
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For more information about California galls, see Ron Russo's
Plant Galls of the Western United States (2021), an update to and expansion of Ron's 2006
Field Guide to Plant Galls of California and Other Western States.
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